贺小彦，博士，副教授，硕士研究生导师

　　邮箱：blsj2009@163.com

　　个人履历

　　2004.09-2008.07：周口师范学院，生物科学专业，学士

　　2008.09-2011.06：福建农林大学，生物化学与分子生物学，硕士

　　2011.09-2015.09：浙江大学，作物栽培学与耕作学，博士

　　2015.10-2018.08：浙江大学，农业与生物技术学院，博士后

　　2017.03-2018.03：澳大利亚Murdoch University，访问学者

　　2018.10-2025.11：青岛农业大学，农学院，讲师

　　2025.12-至今：青岛农业大学，农学院，副教授

　　研究方向：麦类作物作物抗逆基因挖掘及功能研究

　　科研成果

　　一、主持或参与项目

　　1.山东省重点研发计划（农业良种工程项目）子课题，2023LZGC009-2-2,小麦耐盐种质创制和耐盐基因发掘，2023/10-2026/09，主持

　　2.山东省自然科学基金面上项目，ZR2021MC025，大麦抗旱基因HvFRS1的优异等位变异挖掘及转录调控机理解析，2022/01-2024/12，主持

　　3.青岛农业大学高层次人才科研基金项目，663-1119013，大麦抗干旱基因挖掘及功能研究，2019/09-2025/8，主持

　　4.中国博士后基金，2016M590541，西藏野生大麦根毛生长相关基因HvEXPB7的功能鉴定，2016/03-2018/10，主持

　　5.国家自然科学基金青年基金项目，31601231，HvEXPB7介导的西藏野生大麦耐旱机理研究，2017/01-2019/12，主持

　　6.横向课题，作物植株农艺性状与盐离子测定，2024/11-2025/12，主持

　　7.国家自然科学基金面上项目，32372167，tae-miR9774靶向TaDEAH28调控小麦耐盐性的分子机制解析，2024/01-2027/12，参与

　　8.国家自然科学基金面上项目，32171976，大麦D型周期蛋白基因HvCYCD1调控根毛发育的分子机制解析，2022/01-2025/12，参与

　　9.山东省重点研发计划（农业良种工程项目）课题，2021LZGC013-5，特色营养小麦突破性新种质的创制，2021-12至2025-06，参与

　　10.山东省重点研发计划（农业良种工程项目）课题，2023LZGCQY011-2，特色专用小麦新品种选育与配套栽培技术优化及示范推广，2023-09至2026-08，参与

　　二、发表论文

　　1.He, XY., Wang, YJ., Gao, Y., Han, Z., Yin, HY., Zeng, JB., et al. (2025). tae-miR9668 negatively regulates TaRPM1-1A to enhance salt tolerance in wheat, Crop J. 13(6), 1746-1761.

　　2.He, XY., Su, CJ., Zhang, XY., Shi, ZX., Wang, YJ., Peng, HD., et al. (2024). Identification of crucial drought-tolerant genes of barley through comparative transcriptomic analysis and yeast-based stress assay. Front. Genet. 15, 1524118.

　　3.He, XY., He, YR., Dong, YH., Gao, Y., Sun, XC., Chen, WY., et al. (2024). Genome-wide analysis of FRF gene family and functional identification of HvFRF9 under drought stress in barley. Front. Plant Sci. 15, 1347842.

　　4.He, XY., Chen, WY., Sun, XC., Gao, Y., He, YR., Xu, XT., et al. (2023). Genome-wide identification and characterization of glutaredoxin family genes in common wheat. Agronomy 13, 2985.

　　5.He, XY., Han, Z., Yin, HY., Chen, F., Dong, YH., Zhang, LF., et al. (2021). High-throughput sequencing-based identification of miRNAs and their target mRNAs in wheat variety Qing Mai 6 under salt stress condition. Front. Genet. 12, 724527.

　　6.He, XY., Richmond, MEA., Williams, DV., Zheng, WT., and Wu, FB. (2019). Exogenous glycinebetaine reduces cadmium uptake and mitigates cadmium toxicity in two tobacco ggenotypes differing in cadmium tolerance. Int. J. Mol. Sci. 20, 1612.

　　7.He, XY., Zheng, WT., Cao, FB., and Wu, FB. (2016). Identification and comparative analysis of the microRNA transcriptome in roots of two contrasting tobacco genotypes in response to cadmium stress. Sci. Rep. 6, 32805.

　　8.He, XY., Zeng, JB., Cao, FB., Ahmed, IM., Zhang, GP., Vincze, E., et al. (2015). HvEXPB7, a novel β-expansin gene revealed by the root hair transcriptome of Tibetan wild barley, improves root hair growth under drought stress. J. Exp. Bot 66, 7405–7419.

　　9.贺小彦, 王永梅, 何东, 吴港, 孙庆译, 孙雪莲, 刘文星, 曾建斌. (2024) 比较转录组发掘控制大麦根毛长度的基因. 浙江大学学报（农业与生命科学版）, 网络首发.

　　10.Wu G., Sun, XL., Sun, QY., Kang, X., Wang, JY., He, XY., Liu, WX., Xu, DA., Dai, XH., M,WJ., Zeng, JB. (2025) Genetic Variation in Wheat Root Transcriptome Responses to Salinity: A Comparative Study of Tolerant and Sensitive Genotypes. Int. J. Mol. Sci. 6(1), 331.

　　11.Yang, F.H., Sun, X.L., Wu, G., He, X.Y., Liu, W.X., Wang, Y.M., Sun, Q.Y., Zhao, Y., Xu, D.A., Dai, X.H., Ma, W.J., Zeng, J.B.# (2024). Genome-wide identification and expression profiling of the ABF transcription factor family in wheat (Triticum aestivum L.). Int. J. Mol. Sci., 25(7), 3783.

　　12.Zeng, JB., Wang, YM., Wu, G., Sun, QY., He, XY., Zhang, XY., et al. (2024). Comparative Transcriptome Analysis Reveals the Genes and Pathways Related to Wheat Root Hair Length. Int. J. Mol. Sci. 25, 2069.

　　13.Feng, X., Yu, Q., Zeng, JB., He, XY., Ma, WJ., Ge, L., et al. (2023). Comprehensive Analysis of the INDETERMINATE DOMAIN (IDD) Gene Family and Their Response to Abiotic Stress in Zea mays. Int. J. Mol. Sci. 24, 6185.

　　14.Feng, X., Yu, Q., Zeng, JB., He, XY., and Liu, WX. (2022). Genome-wide identification and characterization of GATA family genes in wheat. BMC Plant Biol 22, 372.

　　15.Cheng, CY., Kan, J., Li, S., Jiang, C., He, XY., Shen, H., et al. (2022). Mutation of barley HvPDIL5-1 improves resistance to yellow mosaic virus disease without growth or yield penalties. Front. Plant Sci. 13, 1018379.

　　16.Yin, HY., Sun, Q., Lu, XQ., Zhang, LF., Yuan, YC., Gong, CL., He, XY., et al. (2022a). Identification of the glutamine synthetase (GS) gene family in four wheat species and functional analysis of Ta4D.GSe in Arabidopsis thaliana. Plant Mol Biol. 110, 93–106.

　　17.Yin, HY., Yang, F., He, XY., Du, XY., Mu, P., and Ma, WJ. (2022b). Advances in the functional study of glutamine synthetase in plant abiotic stress tolerance response. Crop J. 10, 917–923.

　　18.Han, Y., Broughton, S., Liu, L., Zhang, XQ., Zeng, JB., He, XY., et al. (2021). Highly efficient and genotype-independent barley gene editing based on anther culture. Plant Commun. 2, 100082.

　　19.Qiu, CW., Liu, L., Feng, X., Hao, PF., He, XY., Cao, FB., et al. (2020). Genome-Wide Identification and Characterization of Drought Stress Responsive microRNAs in Tibetan Wild Barley. Int. J. Mol. Sci. 21, 2795.

　　20.Zeng, JB., Ye, ZL., He, XY., and Zhang, GP. (2019). Identification of microRNAs and their targets responding to low-potassium stress in two barley genotypes differing in low-K tolerance. J. Plant Physiol. 234–235, 44–53.

　　21.Zeng, JB., Quan, XY., He, XY., Cai, SG., Ye, ZL., Chen, G., et al. (2018). Root and leaf metabolite profiles analysis reveals the adaptive strategies to low potassium stress in barley. BMC Plant Biol. 18, 187.

　　22.Ahmed, I. M., Nadira, U. A., Cao, F., He, XY., Zhang, G., and Wu, F. (2016). Physiological and molecular analysis on root growth associated with the tolerance to aluminum and drought individual and combined in Tibetan wild and cultivated barley. Planta 243, 973–985.

　　23.Ahmed, I. M., Nadira, U. A., Bibi, N., Cao, F., He, XY., Zhang, G., et al. (2015). Secondary metabolism and antioxidants are involved in the tolerance to drought and salinity, separately and combined, in Tibetan wild barley. Environ. Exp. Bot. 111, 1–12.

　　24.Cao, F., Cai, Y., Liu, L., Zhang, M., He, XY., Zhang, G., et al. (2015). Differences in photosynthesis, yield and grain cadmium accumulation as affected by exogenous cadmium and glutathione in the two rice genotypes. Plant Growth Regul. 75, 715–723.

　　25.Wang, N., Zhao, J., He, XY., Sun, H., Zhang, G., and Wu, F. (2015). Comparative proteomic analysis of drought tolerance in the two contrasting Tibetan wild genotypes and cultivated genotype. BMC Genomics 16, 432.

　　26.Zeng, J., He, XY., Quan, X., Cai, S., Han, Y., Nadira, U. A., et al. (2015). Identification of the proteins associated with low potassium tolerance in cultivated and Tibetan wild barley. J. Proteomics 126, 1–11.

　　27.Zeng, J., He, XY., Wu, D., Zhu, B., Cai, S., Nadira, U. A., et al. (2014). Comparative Transcriptome Profiling of Two Tibetan Wild Barley Genotypes in Responses to Low Potassium. PLoS ONE 9, e100567.

　　28.Sun, H., Zhang, X., He, XY., Ahmed, I. M., Cao, F., Zhang, G., et al. (2014). N-acetyl-cysteine alleviates Cd toxicity and reduces Cd uptake in the two barley genotypes differing in Cd tolerance. Plant Growth Regul. 74, 93–105.

　　29.Xie, L., He, XY., Shang, S., Zheng, W., Liu, W., Zhang, G., et al. (2014). Comparative proteomic analysis of two tobacco (Nicotiana tabacum) genotypes differing in Cd tolerance. Biometals 27, 1277–1289.

　　30.Dai, H., Ibrahim, W., Zheng, W.-T., Dawood, M., He, XY., Zhao, J., et al. (2013). Characteristics of Photosynthetic Performance, Antioxidant Capacity and Nutrient Concentration of Tibetan Wild Barley in Response to Aluminium Stress. Asian J. Chem. 25, 7727–7731.

　　三、授权专利

　　1.贺小彦; 石兆霞; 穆平; 曾建斌; 尹华燕. TaGRX53在提高植物耐盐性中的应用, 2025-10-21, 中国发明专利, ZL202510748028.1 （第一完成人）

　　2.贺小彦; 张馨怡; 张彩明; 曾建斌; 尹华燕; 穆平. 小麦基因在调控植物耐旱性中的用途, 2026-01-13, 中国发明专利, ZL202511036871.3（第一完成人）

　　3.贺小彦; 穆平; 董义焕; 曾建斌. 一种大麦HvFRF1基因、蛋白、表达载体以及用途, 2024-01-18, 中国发明专利, ZL202211417341.X（第一完成人）

　　4.贺小彦; 曾建斌; 邬飞波. 青藏高原一年生野生大麦HvEXPB7基因及其用途, 2018.01, 中国发明专利, ZL201510494025.6（第一完成人）

　　5.贺小彦; 董义焕; 曾建斌; 尹华燕; 穆平. 一种小麦育种施肥装置, 2021-09-14, 实用新型专利, ZL202023030019.0（第一完成人）​​​​​​​

　　学术兼职情况:

　　山东省科技特派员

　　研究生培养质量情况：

　　2020年开始招收硕士研究生